Lecture 16
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Oldest sedimentary rock
3.86 billion years ago
Stable isotope record
Isotope fractionation
____ isotope is incorporated preferentially over heavy isotope
Lighter
Biomarkers
Molecules known only to be produced biologically
Stromatolites
Microbial fossils Laminated layer of filamentous microorganisms and sediment
Ancient stromatolies associated with light carbon and
2-methylhopanoids
Early life
3.5-2.5 bya
Ancient earth atmosphere was highly ____
Reducing (Fe2)
Chemical disequilibrium between hydrothermal fluid and ocean water
Potential to gain energy
RNA may have been the first
Self-replicating molecule
Self catalytic RNAs may have synthesized proteins, then eventually DNA ____ RNA as the repository for coding information
replaced
Evidence for ____ of ____ becomes evident 2.7-2.4 bya Reduced iron reacted with oxygen to create BIFs which are a major source of iron for mining
oxidation of iron
What created oxygen
oxygenic photosynthesis (splitting water to form H2 and O2) by Cyanobacteria
O2 reacted with UV to produce
Ozone shield
Produced energy for complex life (aerobes) through
Thermodynamic disequilibrium
Endosymbiotic events Mitochondria and chloroplast arose from ____ ____ of prokaryotes within another cell
Symbiotic association
The evolutionary history of a roup of organisms
Phylogenetics
The science of identification, classification, and nomenclature
Taxonomy
Early attempts to understand phylogenetics led to the
Five Kingdom System Based on Morphology Highly subjective
Molecular Evolution
The process of the evolution of DNA, RNA, and proteins
Small subunit rRNA-based phylogeny championed by
Carl Woese
Phylogenetics rRNA genes are perfect for organismal phylogenetics because they are
Universally distributed, functionally homologous, and slow evolving
Phylogenetics Immediately led to discovery of ____, collapse of the five kingdom system, and first meaningful
Archeae/Universal Tree of Life
Phylogeny Why use rRNA
Present in all organisms Structurally and functionally conserved Contains both conserved and variable regions